Our Research
The collections at Rancho La Brea are still at the core of late Pleistocene North American research today.
Our staff, professional paleontologists, and graduate students frequent the collections throughout the year to study a range of topics. Many questions still remain to be answered. If you are interested in using our specimens for research, please visit the Our Collections page.
Collaborative Research Projects
Large multi-institutional projects are exciting opportunities to bring experts from different disciplines together to help answer some of our pressing research questions.
Dr. Jacquelyn Gill, Assistant Professor, Climate Change Institute & School of Biology & Ecology, U. of Maine-Orono
Dr. Jessica Blois, Assistant Professor, Life and Environmental Sciences, UC Merced
Dr. Justin Yeakel, Assistant Professor, Life and Environmental Sciences, UC Merced
We are working on an NSF-funded project to build multi-trophic food webs for La Brea paleo-ecosystems and reconstruct changes in the composition and structure of the ecological network in California across the late 50,000 years.
Using the Project 23 deposits, which span from >50,000 years to ca. 30,000 years ago, we will first reconstruct trophic interactions of the large to small mammals and vegetation from different time slices, allowing us to constrain the range of natural variability among species interactions.
We will then examine the stability of food webs to external perturbations, both simulated (e.g., targeted removal of a species vs. climate change that affects all species individualistically) and observed (e.g., by testing model predictions against actual events in the paleorecord, such as the late Pleistocene megafaunal extinctions). This will give us an idea of how resilient different species were to climate and human impacts and whether properties of the overall interaction networks predict the vulnerability of individual species. This project also involves La Brea curators and staff, and Dr. John Southon at UC Irvine. Stay up-to-date with this project on Twitter at #LaBreaWebs and on our blog.
Dr. Wendy J. Binder, Professor, Loyola Marymount University, CA
Dr. Julie A. Meachen, Assistant Professor, Des Moines University, IA
Dr. F. Robin O’Keefe, Professor, Marshall University, WV
Dr. Larisa DeSantis, Associate Professor, Vanderbilt University, TN
Dr. Emily Lindsey, Assistant Curator, La Brea Tar Pits Museum, CA
Dr. John Southon, Researcher, Earth System Science, UC Irvine, CA
The La Brea fossils span a critical time in Earth’s history (approximately 50,000 years ago to the present) that includes major events such as the end of the last Ice Age, the arrival of humans in North America, and an extinction that killed two thirds of the large mammals on the continent.
This unique deposit allows us to investigate relationships between major environmental changes and evolutionary variation (size, diet, etc.) in large mammals, information that is critical for promoting the survival of wildlife today. There is a distinct lack of radiocarbon dates for fossils in this period, resulting in our inability to correlate biotic change with time.
This project will intensively radiocarbon date samples from multiple pits spanning the last 50,000 years to establish the first detailed chronology for the entrapment of four extinct species of large mammals (saber-toothed cats, dire wolves, Antique bison, and the Western horse), and one extant species (the coyote). It will involve the collection and compilation of census data for these species to track changes in total abundance and diversity, as well as data on morphological and dietary changes. A well-resolved chronology will allow these data on evolutionary changes in mammals and the Pleistocene megafaunal extinctions to be linked to existing records of major paleo-environmental changes. This work will have broad implications for our understanding of extinctions, survival, environmental variables, and humans on mammalian ecology, which is directly relevant to modern conservation. The project also involves community outreach in the greater Los Angeles area through the La Brea Tar Pits Museum and will create educational content that will be available online for high school teachers across the country.
Dr. Benjamin Nye, Director for Learning Science Research, Institute for Creative Technologies, University of Southern California, CA
Dr. Emily Lindsey, Assistant Curator, La Brea Tar Pits Museum, CA
Dr. Gale M. Sinatra, Professor, Rossier School of Education, University of Southern California, CA
Dr. William Swartout, Chief Technology Officer, University of Southern California, CA
Re-Living Paleontology is an AISL Research in Service to Practice project to study how visual immersion and interactivity in augmented reality (AR) affects visitors' engagement and understanding of science. This project will communicate paleontology research from the La Brea Tar Pits to the general public (e.g., adults, families) through an AR-enhanced timeline exhibit in the public park next to the museum. Currently, many informal learning settings are considering AR to raise engagement and communicate science. To achieve that potential, studies are needed to demonstrate when and how to leverage AR so that it increases knowledge rather than merely entertains visitors. Unfortunately, studies on AR for informal learning typically foreclose on high-level design choices at the formative stages, so empirical results on learner outcomes are seldom reported. This project addresses these gaps through a user-centered design and evaluation process with both formative and comparative studies.
This project investigates two high-level design factors for mobile AR: visual immersion and interactivity. These impact the learning experience and the development so extensively that multiple versions are seldom compared. These factors also have unique considerations for informal settings, such as how to balance immersion against situational awareness (e.g., 3D viewers reduce field of view). One goal of this project is to systematically compare qualitatively different AR designs that convey equivalent science content, so that we can study these tradeoffs empirically. The second goal is to leverage these findings to publicly release an AR experience that promotes engagement, increases understanding of science, and reduces scientific misconceptions. In that experience, visitors engage with the scientific process by tracing research questions that have been studied at La Brea. At waypoints on the timeline, visitors will explore AR time portals where they gather evidence to distinguish between competing hypotheses and update their hypotheses as they find new evidence.
This work is important because many U.S. citizens misunderstand and question claims that have wide support among scientists, have misconceptions about critical issues, and lack a basic appreciation of how scientists evaluate evidence. Public understanding of science affects critical individual decisions and public policy. Informal learning institutions offer powerful settings to make a lasting impact on building knowledge and increasing engagement with science: a single exhibit can reach hundreds of thousands of visitors. AR is positioned to play a key role in the future of such high-impact exhibits, since it supports immersive, personalized interactions with hidden structural and causal relationships in situ. This project will help achieve that promise by studying how AR design factors affect visitor outcomes.
So much still to learn!
Today's research ranges from traditional taxonomic descriptions to addressing functional morphology questions to investigating how ecosystems have been altered under changing climatic conditions over the past 50,000 years in the Los Angeles region. The ability to directly date our fossils using the unstable isotope carbon-14 enables us to pair these results with cutting-edge techniques such as stable isotopes, dental microwear, and collagen sequencing to help us understand the factors that influenced the current distributions of much of our flora and fauna before humans changed the landscape. This provides a deeper time perspective by which we can understand the past and present, as well as anticipate future ecosystems in order to support their resilience. Hundreds of publications have resulted from many years of research. You can find these citations below split into broad categories. You can also download La Brea and Beyond: The Paleontology of Asphalt-Preserved Biotas for free!
- Bromage, T. G. & Shermis, S. 1981. The La Brea Woman (HC 1323): Descriptive analysis. Society of California Archaeology, Occasional Papers, 3, 59-75.
- Eberhart, H. 1961. The cogged stones of Southern California. American Antiquity, 26 (3), 361-370.
- Haynes, G. & Sanford, D. 1984. On the possible utilization of Camelops by early man in North America. Quaternary Research, 22, 216-230.
- Heizer, R.F. 1953. Aboriginal use of bitumen by the California Indians. Bulletin of the California Division of Mines, 118, 74.
- Kennedy, G. E. 1989. A note on the ontogenetic age of the Rancho La Brea hominid, Los Angeles, California. Bulletin of the Southern California Academy of Sciences, 88 (3), 123-126.
- Kuzminsky, S. C. 2012. Rancho La Brea Woman: a new 3D analysis of a 9,000-year-old Paleoamerican cranium from southern California. In American Journal of Physical Anthropology, Vol. 147, pp. 187-187.
- Merriam, J. C. 1914. Preliminary report on the discovery of human remains in an asphalt deposit at Rancho La Brea. Science, 40, 198-203.
- Merriam, J. C. 1914. The brea maid. Bulletin of the Southern California Academy of Sciences, 13 (2), 27-29.
- Miller, G. J. 1975. A study of cuts, grooves, and other marks on recent and fossil bones: II. Weathering cracks, fractures, splinters, and other similar natural phenomena. In E. Swanson (Ed.), Lithic technology, (pp. 211-226). The Hague: Mouton Publishing.
- Miller, G. J. 1969. A study of cuts, grooves, and other marks on recent and fossil bone: I. Animal tooth marks. Idaho State Museum Tebiwa 12 (1), 20-26.
- Miller, G. J. 1969. Man and Smilodon: A preliminary report on their possible coexistence at Rancho La Brea. Los Angeles County Museum of Natural History Contributions in Science, 163, 1-8.
- Reynolds, R. L. 1985. Domestic dog associated with human remains at Rancho La Brea. Bulletin of the Southern California Academy of Sciences, 84 (1), 76-85.
- Salls, R. A. 1991. Ancient brea people. Terra Magazine, Natural History Museum of Los Angeles County, 31 (1), 10-11.
- Salls, R. A. 1986. The La Brea atlatl foreshafts: Inferences for the Millingstone Horizon. Pacific Coast Archaeological Society Quarterly, 22 (2), 21-30.
- Salls, R. A. 1980. The La Brea cogged stone. Masterkey, Southwest Museum, 54 (2), 53-59.
- Tate, B. 1995. Evidence for atlatls at Rancho La Brea. The Atlatl: The Newsletter of the World Atlatl Association, Incorporated, 8 (4), 1-3.
- Wärmländer, S. K. T. S., S. B. Sholts, J. M. Erlandsen, T. Gjerdrim, and R. Westerholm. 2011. Could the health decline of prehistoric California Indians be related to exposure to polycyclic aromatic hydrocarbons (PAHs) from natural bitumen. Environmental Health Perspectives
- Woodard, A. 1937. Atlatl dart foreshafts from the La Brea pits. Bulletin of the Southern California Academy of Sciences, 36 (2), 41-59.
- Bochenski, Z. & Campbell, K. E. 2006. The extinct California turkey, Meleagris californica, from Rancho La Brea: Comparative osteology and systematics. Los Angeles County Museum of Natural History Contributions in Science, 509, 1-92.
- Campbell, K. E. 2002. A new species of late Pleistocene lapwing from Rancho La Brea, California. The Condor, 104(1), 170-174.
- Campbell, K. E. 1995. A review of the cranes (Aves: Gruidae) of Rancho La Brea, with the description of a new species. Los Angeles County Museum of Natural History Contributions in Science, 452, 1-13. [TYPE: Grus paei sp. nov.]
- Campbell, K. E. & Bochenski, Z. M. 2015. The Owls (Aves: Strigiformes) of Rancho La Brea. La Brea and beyond: The paleontology of asphalt-preserved biotas, ed. JM Harris. Natural History Museum of Los Angeles County, Science Series, (42), 5-21.
- Campbell K. E. & Bochenski Z. 2012. Two new late Pleistocene miniature owls from Rancho La Brea, California. Acta Palaeontologica Polonica 58 (4), 2013: 707-721 doi: http://dx.doi.org/10.4202/app.2011.0125
- Campbell, K. E. & Bochenski, Z. 2010. A new genus for the extinct late Pleistocene owl Strix brea Howard (Aves: Strigiformes) from Rancho La Brea, California. Records of the Australian Museum, 62, 123-144.
- Compton, L. V. 1934. New bird records from the Pleistocene of Rancho La Brea. In J. Grinnell (Ed.), The Condor, 36 (5), (pp. 221-222). Berkeley: Cooper Ornithological Club.
- Dawson, W. R. 1948. Records of fringillids from the Pleistocene of Rancho La Brea. In A. H. Miller (Ed.), The Condor, 50 (2), (pp. 57-63). Berkeley: Cooper Ornithological Club.
- Engles, W. L. 1935. Status of Toxostoma redivivum in the Rancho La Brea fauna. In J. Grinnell (Ed.), The Condor, 37 (6), (p. 258). Berkeley: Cooper Ornithological Club.
- Howard, H. 1974. Postcranial elements of the extinct condor Breagyps clarki (Miller). Los Angeles County Museum of Natural History Contributions in Science, 256, 1-24.
- Howard, H. 1972. The incredible Teratorn again. The Condor, 74 (3), 341-344.
- Howard, H. 1972. Type specimens in the collections of the Natural History Museum of Los Angeles County. Los Angeles County Museum of Natural History Contributions in Science, 228, 1-27.
- Howard, H. 1968. Limb measurements of the extinct vulture, Coragyps occidentalis. Papers of the Archaeological Society of New Mexico, 1, 115-128.
- Howard, H. 1964. A fossil owl from Santa Rosa Island, California, with comments on the eared owls of Rancho La Brea. Bulletin of the Southern California Academy of Sciences, 63 (1), 27-31.
- Howard, H. 1962. Fossil birds, with especial reference to the birds of Rancho La Brea (Revised edition). Los Angeles County Museum of Natural History, Science Series no.17, Paleontology no.10, (February 1962), 3-44.
- Howard, H. 1955. Fossil birds, with special reference to the birds of Rancho La Brea. Los Angeles County Museum, Science Series 17, Paleontology, 10, 3-40.
- Howard, H. 1950. Teratornis: Wonder bird of the Ice Age. Los Angeles County Museum Leaflet Series, Science, No. 3 (Nov. 1950), 1-4.
- Howard, H. 1947. California’s flightless birds. Los Angeles County Museum Quarterly, 6 (2), 7-11.
- Howard, H. 1947. An ancestral Golden Eagle raises a question in taxonomy. The Auk, 64, 287-291.
- Howard, H. 1945. Fossil birds. Los Angeles County Museum, Science Series, No. 10, Paleontology, 6, 1-40.
- Howard, H. 1945. Observations on young tarsometatarsi of the fossil turkey Parapavo californicus (Miller). The Auk, 62, 596-603.
- Howard, H. 1942. A review of the American fossil storks. Carnegie Institution of Washington Publications, 530 (7), 187-203.
- Howard, H. 1938. The Rancho La Brea caracara: A new species. [TYPE: Polyborus prelutosus sp. nov.] Carnegie Institution of Washington Publications, 487 (5), 217-240.
- Howard, H. 1937. A Pleistocene record of the passenger pigeon in California. In J. Grinnell (Ed.), The Condor, 39 (1), (pp. 12-14). Berkeley: Cooper Ornithological Club.
- Howard, H. 1936. Further studies upon the birds of the Pleistocene of Rancho La Brea. In J. Grinnell (Ed.), The Condor, 38 (1), (pp. 32-36). Berkeley: Cooper Ornithological Club.
- Howard, H. 1935. The Rancho La Brea wood ibis. [TYPE: Mycteria wetmorei sp. nov.]. The Condor, Vol. 37, No. 5 (Sep. - Oct., 1935), pp. 251-253
- Howard, H. 1933. A new species of owl from the Pleistocene of Rancho La Brea, California. [TYPE: Strix brea sp. nov.] In J. Grinnell (Ed.), The Condor, 35 (2), (pp. 66-69). Berkeley: Cooper Ornithological Club.
- Howard, H. 1932. Eagles and eagle-like vultures of the Pleistocene of Rancho La Brea. Carnegie Institution of Washington Publications, 429, 1-82.
- Howard, H. 1930. A census of the Pleistocene birds of Rancho La Brea from the collection of the Los Angeles Museum. In J. Grinnell (Ed.), The Condor, 32 (2), (pp. 81-88). Berkeley: Cooper Ornithological Club.
- Howard, H. 1929. Additional bird records from the Pleistocene of Rancho La Brea. In J. Grinnell (Ed.), The Condor, 31 (6), (pp. 251-252). Berkeley: Cooper Ornithological Club.
- Howard, H. 1928. The beak of Parapavo californicus (Miller). Bulletin of the Southern California Academy of Sciences, 27 (3), 90-91.
- Howard, H. 1927. A review of the fossil bird, Parapavo californicus (Miller), from the Pleistocene asphalt beds of Rancho La Brea. University of California Publications, Bulletin of the Department of Geological Sciences, 17 (1). In G. D. Louderback, & W. D. Matthew (Eds.), University of California Publications in Geological Sciences, (pp. 1-62). Berkeley: University of California Press.
- Howard, H. & Miller, A. H. 1939. The avifauna associated with the human remains at Rancho La Brea, California. Carnegie Institution of Washington Publications, 514, 39-48.
- Husband, R. A. 1924. Variability in Bubo virginianus from Rancho La Brea. In J. Grinnell (Ed.), The Condor, 26 (6), (pp. 220-225). Berkeley: Cooper Ornithological Club.
- Miller, A. H. 1947. A new genus of icterid from Rancho La Brea. [TYPE: Pandanaris convexa gen. et sp. nov.]. In A. H. Miller (Ed.), The Condor, 49 (1), (pp. 22-24). Berkeley: Cooper Ornithological Club.
- Miller, A. H. 1929. Additions to the Rancho La Brea avifauna. In J. Grinnell (Ed.), The Condor, 31 (5), (pp. 223-224). Berkeley: Cooper Ornithological Club.
- Miller, A. H. 1929. The passerine remains from Rancho La Brea in the paleontological collection of the University of California. University of California Publications, Bulletin of the Department of Geological Sciences, 19 (1). In G. D. Louderback (Ed.), University of California Publications in Geological Sciences, 1929-1931, (pp. 1-22). Berkeley: University of California Press.
- Miller, L.H. 1938. A study of the skull of the Pleistocene stork, Ciconia maltha, Miller. Transactions of the San Diego Society of Natural History, 3 (4), 455-462.
- Miller, L. H. 1924. Anomalies in the distribution of fossil gulls. In J. Grinnell (Ed.), The Condor, 26 (5), (pp.173-174). Berkeley: Cooper Ornithological Club.
- Miller, L. H. 1916. The owl remains from Rancho La Brea. University of California Publications, Bulletin of the Department of Geology/University of California Publications in Geology, 9 (8). In A. C. Lawson, & J. C. Merriam (Eds.), University of California Publications in Geological Sciences, 1915-1916, (pp. 97-104). Berkeley: University of California Press.
- Miller, L. H. 1916. Two vulturid raptors from the Pleistocene of Rancho La Brea. [TYPE: Neophrontops americanus gen. et sp. nov.; Neogyps errans gen. et sp. nov.] University of California Publications, Bulletin of the Department of Geology/University of California Publications in Geology, 9 (9). In A. C. Lawson, & J. C. Merriam (Eds.), University of California Publications in Geological Sciences, 1915-1916, (pp. 105-109). Berkeley: University of California Press.
- Miller, L. H. 1916. A review of the species Pavo californicus. [TYPE: Parapavo californicus gen. nov.] University of California Publications, Bulletin of the Department of Geology/University of California Publications in Geology, 9 (7). In A. C. Lawson, & J. C. Merriam (Eds.), University of California Publications in Geological Sciences, 1915-1916, (pp. 89-96). Berkeley: University of California Press.
- Miller. L. H. 1915. A walking eagle from Rancho La Brea. [TYPE: Morphnus daggetti sp. nov.]. In J. Grinnell (Ed.), The Condor, 17 (5), (pp. 179-181). Hollywood, CA: Cooper Ornithological Club.
- Miller, L. H. 1911. A series of eagle tarsi from the Pleistocene of Rancho La Brea. University of California Publications, Bulletin of the Department of Geology. , 6 (12). In A. C. Lawson, & J. C. Merriam (Eds.), University of California Publications in Geological Sciences, 1910-1911, (pp. 305-316). Berkeley: University of California Press.
- Miller, L. H. 1910. Wading birds from the Quaternary asphalt beds of Rancho La Brea. University of California Publications, Bulletin of the Department of Geology, 5 (30). In A. C. Lawson (Ed.), University of California Publications in Geological Sciences, 1906-1910, (pp. 439-448). Berkeley: University of California Press.
- Miller, L. H. 1910. The condor-like vultures of Rancho La Brea. [TYPE: Sarcorhamphus clarki sp. nov.; Pleistogyps rex gen. nov.; Cathortornis gracilis sp. nov.]. University of California Publications, Bulletin of the Department of Geology, 6 (1). In A. C. Lawson, & J. C. Merriam (Eds.), University of California Publications in Geological Sciences,1910-1911, (pp. 1-19). Berkeley: University of California Press.
- Miller, L. H. 1910. Fossil birds from the Quaternary of Southern California. In J. Grinnell (Ed.), The Condor, 12 (1), (pp. 12-15). Hollywood, CA: Cooper Ornithological Club of California.
- Miller, L. H. 1909. Teratornis, a new avian genus from Rancho La Brea. University of California Publications, Bulletin of the Department of Geology, 5 (21). In A. C. Lawson (Ed.), University of California Publications in Geological Sciences, 1906-1910, (pp. 305-317). Berkeley: University of California Press.
- Miller, L. H. 1909. Pavo californicus, a fossil peacock from the Quaternary asphalt beds of Rancho La Brea. University of California Publications, Bulletin of the Department of Geology, 5 (19). In A. C. Lawson (Ed.), University of California Publications in Geological Sciences, 1906-1910, (pp. 285-289). Berkeley: University of California Press.
- Miller, L. H. & Howard, H. 1938. The status of the extinct condor-like birds of the Rancho La Brea Pleistocene. Publications of the University of California, Los Angeles in Biological Sciences, 1 (9), 169-176.
- Olson, S. L. 2007. The “Walking Eagle” Wetmoregyps daggetti Miller: a scaled-up version of the Savanna Hawk (Buteogallus meridionalis). Ornithological Monographs No. 63: 110-114.
- Sibley, C. 1939. Chipping sparrows in the Rancho La Brea. The Condor, 41 (6), 258-259. Berkeley, CA: Cooper Ornithological Club.
- Sibley, C. 1939. Fossil fringillides from Rancho La Brea. The Condor, 41 (3), 126-127. Berkeley, CA: Cooper Ornithological Club.
- Sushkin, P. P. 1928. On the affinities of Pavo californicus. Ibis, January, 135-138.
- Akersten, W. A., Foppe, T. M., & Jefferson, G. T. 1988. New source of dietary data for extinct herbivores. Quaternary Research, 30, 92-97. The University of Washington.
- Axelrod, D. I. 1981. Holocene climatic changes in relation to vegetation disjunction and speciation. The American Naturalist 117 (6): 847-870.
- Davidson, A. 1914. The oldest known tree. Bulletin of the Southern California Academy of Sciences, 13 (1), 14-16.
- Frost, F. H. 1927. The Pleistocene flora of Rancho La Brea. University of California Publications in Botany, 14 (3), 73-98.
- Gerhart, L. M., Harris, J.M., Nippert, J. B., Sandquist D.R. & Ward J.K. 2012. Glacial trees from the La Brea tar pits show physiological constraints of low CO2. New Phytologist. 194(1):63-9.
- George, J., Dimson, M., Dunn, R.E., Lindsey, E.L., Farrell, A.B., Paola Aguilar, B., & MacDonald, G.M. 2024. Identification of fossil juniper seeds from Rancho La Brea (California, USA): drought and extirpation in the Late Pleistocene. New Phytologist. https://doi.org/10.1111/nph.20324
- Heusser, L.E., 1998. Direct correlation of millennial-scale changes in western North America vegetation and climate with changes in the California Current system over the past ~60 kyr. Paleoceanography 13, 252-262.
- Knowlton, F. H. 1916. Notes on two conifers from the Pleistocene Rancho La Brea asphalt deposits near Los Angeles, California. Journal of the Washington Academy of Science, 6, 85-86.
- Li, G., Gerhart, L. M., Harrison, S. P., Ward, J. K., Harris, J. M., & Prentice, I. C. 2017. Changes in biomass allocation buffer low CO2 effects on tree growth during the last glaciation. Scientific Reports, 7.
- Mason, H. L. 1927. Fossil records of some west American conifers. Carnegie Institution of Washington Publications, 346 (5), 139-160.
- Ney, R. W. C., and E. C. Jeffrey. 1922. Flora of the Rancho La Brea. Bulletin of the Geological Society of America, 33, 204.
- Templeton, B. C. 1964. The fruits and seeds of the Rancho La Brea Pleistocene deposits. Doctoral Dissertation, Oregon State University, Corvallis, Oregon.
- Templeton, B. C. 1955. Fossil plants in the La Brea deposits. Los Angeles County Museum of Natural History Quarterly, 12 (1), 8-11.
- Ward, J, K., Harris, J. K., Cerling, T. E., Weidenhoeft, A., Lott, M. J., Dearing, M., Coltrain J. B. & Ehleringer J.R. 2005. Carbon starvation in glacial trees recovered from the La Brea tar pits, southern California. Proceedings of the National Academy of Sciences, 102, 690-694.
- Warter, J. K. 1976. Late Pleistocene plant communities – Evidence from the Rancho La Brea tar pits. Symposium Proceedings on Plant Communities of Southern California, Special Publication of the California Native Plant Society, 2, 32-39.
- Berta, A. 1985. The status of Smilodon in North and South America. Natural History Museum of Los Angeles County Contributions in Science, 370, 1-15.
- Bovard, J. F. 1907. Notes on Quaternary Felidae from California. University of California Publications, Bulletin of the Department of Geology, 5 (10). In A. C. Lawson (Ed.), University of California Publications in Geological Sciences, 1906-1910, (pp. 155-170). Berkeley: University of California Press.
- Cox, S. M. & Jefferson, G. T. 1988. The first individual skeleton of Smilodon from Rancho La Brea. Current Research in the Pleistocene, 5, 66-67.
- Dundas, R. G. 2008. Quaternary records of the dire wolf, Canis dirus, in North and South America. Boreas 28 (3): 375-385.
- Emslie, S. D. & Czaplewski, N. J. 1985. A new record of giant short-faced bear, Arctodus simus, from western North America with a re-evaluation of its paleobiology. Natural History Museum of Los Angeles County Contributions in Science, 371, 1-12.
- Hartstone-Rose, A., Dundas, R. G., Boyde, B., Long, R. C., Farrell, A. B., & Shaw, C. A. 2015. The Bacula of Rancho La Brea. La Brea and beyond: The paleontology of asphalt-preserved biotas, ed. JM Harris. Natural History Museum of Los Angeles County, Science Series, (42), 53-63.
- Hartstone-Rose, A., Long, R. C., Farrell, A. B. & Shaw, C.A. 2012. The clavicles of Smilodon fatalis and Panthera atrox (Mammalia: Felidae) from Rancho La Brea, Los Angeles, California. Journal of Morphology, 273, 981-991.
- Jefferson, G. T. 1983. First record of jaguar from the late Pleistocene of California. Bulletin of the Southern California Academy of Sciences, 82 (2), 95-98.
- Jefferson, G. T. & Tejada-Flores, A. E. 1993. The late Pleistocene record of Homotherium (Felidae: Machariodontinae) in the southwestern United States. PaleoBios, 15 (3), 37-46.
- Kurten, B. 1960. A skull of the grizzly bear (Ursus arctos L.) from Pit 10, Rancho La Brea. Los Angeles County Museum of Natural History Contributions in Science, 39, 1-7.
- Kurtén, B., and L. Werdelin. 2010. Relationships between North and South American Smilodon. Journal of Vertebrate Paleontology 10 (2): 158-169.
- Merriam, J. C. 1918. Note on the systematic position of the wolves of the Canis dirus group. [TYPE: Aenocyon dirus gen. nov.]. University of California Publications, Bulletin of the Department of Geology/University of California Publications in Geology, 10, (27). In A. C. Lawson, & J. C. Merriam (Eds.), University of California Publications in Geological Sciences, 1916-1918, (pp. 531-533). Berkeley: University of California Press.
- Merriam, J. C. 1918. New puma-like cat from Rancho La Brea. [TYPE: Felis daggetti sp. nov.]. University of California Publications, Bulletin of the Department of Geology/University of California Publications in Geology, 10 (28). In A. C. Lawson, & J. C. Merriam (Eds.), University of California Publications in Geological Sciences, 1916-1918, (pp. 535-537). Berkeley: University of California Press.
- Merriam, J. C. 1912. Recent discoveries of Carnivora in the Pleistocene of Rancho La Brea. University of California Publications, Bulletin of the Department of Geology/University of California Publications in Geology, 7 (3). In A. C. Lawson, & J. C. Merriam (Eds.), University of California Publications in Geological Sciences 1912-1924,(pp. 39-46). Berkeley: University of California Press.
- Merriam, J. C. 1912. The fauna of Rancho La Brea: Part 2: Canidae. [TYPE: Canis milleri sp. nov.; Canis ochropus orcutti ssp. nov.]. Memoirs of the University of California, 1 (2), 217-272.
- Merriam, J. C. 1911. Note on a gigantic bear from the Pleistocene of Rancho La Brea. [TYPE: Arctotherium californicum sp. nov.]. University of California Publications, Bulletin of the Department of Geology, 6 (6). In A. C. Lawson, & J. C. Merriam (Eds.), University of California Publications in Geological Sciences, 1910-1911, (pp. 163-166). Berkeley: University of California Press.
- Merriam, J. C. 1910. New mammalia from Rancho La Brea. [TYPE: Canis orcutti sp. nov.; Canis andersoni sp. nov; Canis occidentalis furlongi ssp. nov.; Lynx californicus fisheri ssp. nov.]. University of California Publications, Bulletin of the Department of Geology, 5 (25). In A. C. Lawson (Ed.), University of California Publications in Geological Sciences 1906-1910, (pp, 391-395). Berkeley: University of California Press.
- Merriam, J. C. 1909. The skull and dentition of an extinct cat closely allied to Felis atrox Leidy. [TYPE: Felis atrox bebbi ssp. nov.] University of California Publications, Bulletin of the Department of Geology, 5 (20). In A. C. Lawson (Ed.), University of California Publications in Geological Sciences, 1906-1910), (pp. 291-304). Berkeley: University of California Press.
- Merriam, J. C. & Stock, C. 1932. The Felidae of Rancho La Brea. [TYPE: Felis bituminosa sp. nov.; Smilodon californicus brevipes ssp. nov.]. Carnegie Institution of Washington Publication #422. Washington, D. C.: Carnegie Institution of Washington.
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